
Yoshimura's kakuja kagune also forms a mask similar to the One-Eyed Owl's with minor differences in the eye-hole size and line curves. While his ukaku kagune extends from middle to far distances, his kakuja cuts down those in front it is almost impossible to escape Yoshimura's reach. With two large blades, he can cover the distance between his opponents, parry incoming attacks, and strike back synchronously. Having cannibalized ghouls during his missions to exterminate other ghoul organizations, Yoshimura sharpened and developed his kagune to form two large scimitar blades from his shoulders to counteract this weakness. Ukaku ghouls are known to specialize in long-range attacks and can be overwhelmed at a close-range. It should be noted that members of the Tsukiyama family have a birth defect where cannibalism has little to no chance of triggering this mutation in their Rc cells due to consanguineous marriages, thereby preventing kakuja mutations. Yoshimura's kakuja mask also grew to resemble the One-Eyed Owl's mask, with the former holding the desire to keep attention towards himself and away from the latter for personal reasons.It is explained by Eto that the size of a kagune is determined by rc quantity and quality while the shape of it is determined by imagination and creativity, which also applies to kakujas hence Kaneki's centipede and Yoshimura resembling a one eyed owl. In addition, Kaneki compares himself to Gregor Samsa, the protagonist of the novel The Metamorphosis who turned into an insect-like creature, all while picturing a centipede in his mind. The transformation seems to be affected by psychological factors as Ken Kaneki's kakuja kagune takes the form of a centipede, resembling the Red-headed Chinese Centipede Yakumo Oomori used to torture Kaneki by putting it in his ear. For instance, while the One-Eyed Owl had a human-shape and size before its disappearance, it eventually developed into a much larger and more monstrous form during the Owl Suppression Operation in the 20th ward. If the concentration becomes high enough, it may trigger a mutation and lead to the formation of a new kagune.Ī kakuja can further evolve by continuing to cannibalize their kind. It is said that through cannibalism, a ghoul absorbs Rc cells, and this increases the concentration of Rc cells within a ghoul's body. Sometimes, the transformation is incomplete, and such individuals are called half-kakujas (半赫者, han kakuja).

When a ward has a higher ghoul to human ratio, cannibalism is more common due to turf wars and limited food supply. Though it is rare, it can occur through repeated cannibalism. A Kakuja (赫者, red one, kakuja) is a kind of ghoul with a transformed kagune that clads the ghoul's body.
